Phylogenetic trees are now fundamental in many areas of biology. This course provides as in-depth introduction to phylogenetic biology including theoretical foundations, approaches to phylogenetic tree inference, and applications of phylogenetic trees in ecology and evolution. Students will gain skills in bioinformatics, with application to DNA sequence analysis and phylogenetic tree inference. Through a combination of lectures, discussions, and computer labs, students will master the theory and practice of phylogenetic tree inference and phylogenetic methods in ecology and evolution.